|English||(latin for) Little Man|
The Homunculus (ホムンクルス, Homunkurusu; ; ; ) are a group of artificially created humans who seek to fulfill their leader's goals of creating a Philosopher's Stone. Each homunculus is named after one of the seven deadly sins, and, with the exception of Pride, can be identified by their bearing of the mark of the Ouroboros somewhere on their body. They also possess an immensely heightened regenerative ability, rendering them nigh indestructible, though not invincible. Aside from this, the concept of the homunculi varies drastically between the manga and first anime series.
In the manga, the homunculi were created by or are at the discretion of Father, being a manifestation of one of his traits: his greed, his lust, and so on. Father would later reveal that he ultimately created the homunculi for the sole purpose of purifying himself of his personal flaws.
Each of the homunculi has a Philosopher's Stone somewhere in his or her body which fuels their regenerative ability, and were created at the same time as their respective Philosopher's Stone. While the first six were originally created from Father's own being, both Wrath and the second Greed were humans who accepted a Philosopher's Stone. The only way to kill a homunculus is by expending all of the power in their Philosopher's Stone, leaving them unable to revive themselves once they have been killed.